It's not because you have spark with the spark plugs off that they do fire when bolted in the head.
I know it's may sound stupid, but with the pressure inside the cylinder as well as the air/fuel mixture it's more difficult for the spark to happen.
It did happen to me and I did change the 4 plugs and it fired right away. I'm not saying that this is your problem, but it's a cheap and easy thing to change.
Frank
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