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If you touch the throttle (while O2 sensor is unplugged) and get a lean condition its either
the maf (a boost leak, pressure leak, poor maf placement, insensitive/dirty maf, etc... maf related)
or
the injectors (wrong size, leaking, not leaking, not opening properly (low voltage or w/e), dirty filthy rail or injectors, fuel pressure drop, etc...)
or
It needs tuning (wrong ecu, wrong tune, poor wiring connections, poor grounding, Bad O2 sensor BLM (Long term memory), etc...)
Because it goes like this:
maf -> ecu -> injectors
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