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The FPR keeps the supply line from the pump to the FPR at the set PSI. Your injectors need to be between the Pump and the FPR to be held at pressure.
After the FPR is the return line. Any pressure in this line is just from the resistance of the return line. The stock rail mount FPR's hose connection is the return. The line directly into the rail is the supply.
Maybe I missed it but, What FPR and gauge are you using? Remote or Rail mount?
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Comments should be taken as Opinions not as Statements of Fact 
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